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Megaman X was built by Dr. Light while Megaman was fighting Wily. Wily then one day released his super robot, Zero, to destroy everyone. Zero killed Megaman, Protoman, Roll, and the others. After Megaman got destroyed, Dr. Light released X, who was a far more advanced combat version of Megaman.
Of course all of this is speculation, but chances are most of it is true. Also the X series has a different main enemy (Sigma), all different enemies cept the helmet guys, and the bosses are all completely different (Based of animals, not humanoids). Plus the story is alot different.
As for how Megaman Legends is... its a action "rpg". You do not fight bosses and gain their powers, instead you find weapons like Grenade Launchers and what not. The game is also in full 3d, and has shops and NPC's and the whole works. The game also has a half decent story, and the action is alright. In my eyes not a real Megaman game, but fun nonetheless.
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